The colder temperature does not effect the actual gauge. It does determine what pressure the liquid CO2 in the cylinder exerts. If the cylinder is at ambient outside the refrigerator, the high pressure gauge would read around 825 PSI until all of the liquid is gone, then the pointer begins to drop. Inside the box, around 450 or less depending on the temperature of the box.
The low pressure gauge simply tells you what pressure you have regulated the cylinder pressure down to for use at the kegs.
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